$_ The default input and pattern-searching space. The
following pairs are equivalent:

while (<>) {... # only equivalent in while!
while ($_ = <>) {...

/^Subject:/
$_ =~ /^Subject:/

y/a-z/A-Z/
$_ =~ y/a-z/A-Z/

chop
chop($_)

(Mnemonic: underline is understood in certain opera-
tions.)